{"id":85885,"date":"2026-03-13T15:46:50","date_gmt":"2026-03-13T15:46:50","guid":{"rendered":"https:\/\/green-athens.gr\/?p=85885"},"modified":"2026-03-22T09:05:30","modified_gmt":"2026-03-22T09:05:30","slug":"casuarina-equisetifolia","status":"publish","type":"post","link":"https:\/\/green-athens.gr\/en\/casuarina-equisetifolia\/","title":{"rendered":"Australian pine"},"content":{"rendered":"<section class=\"wpb-content-wrapper\"><p>[vc_row unlock_row_content=&#8221;yes&#8221; row_height_percent=&#8221;100&#8243; override_padding=&#8221;yes&#8221; h_padding=&#8221;3&#8243; top_padding=&#8221;3&#8243; bottom_padding=&#8221;3&#8243; back_image=&#8221;86162&#8243; kburns=&#8221;yes&#8221; overlay_color=&#8221;color-742106&#8243; overlay_alpha=&#8221;80&#8243; overlay_color_blend=&#8221;multiply&#8221; gutter_size=&#8221;3&#8243; column_width_percent=&#8221;100&#8243; shift_y=&#8221;0&#8243; z_index=&#8221;0&#8243; top_divider=&#8221;fan-opacity&#8221; bottom_divider=&#8221;gradient&#8221; changer_back_color=&#8221;yes&#8221; uncode_shortcode_id=&#8221;726788&#8243; overlay_color_type=&#8221;uncode-palette&#8221; row_name=&#8221;title&#8221;][vc_column column_width_percent=&#8221;80&#8243; position_horizontal=&#8221;left&#8221; position_vertical=&#8221;middle&#8221; gutter_size=&#8221;3&#8243; style=&#8221;dark&#8221; font_family=&#8221;font-156269&#8243; overlay_alpha=&#8221;50&#8243; shift_x=&#8221;0&#8243; shift_y=&#8221;0&#8243; shift_y_down=&#8221;0&#8243; z_index=&#8221;0&#8243; align_medium=&#8221;align_left_tablet&#8221; medium_width=&#8221;0&#8243; align_mobile=&#8221;align_left_mobile&#8221; mobile_width=&#8221;0&#8243; zoom_width=&#8221;0&#8243; zoom_height=&#8221;0&#8243; width=&#8221;1\/1&#8243; uncode_shortcode_id=&#8221;183356&#8243;][vc_message css_animation=&#8221;zoom-in&#8221; uncode_shortcode_id=&#8221;166414&#8243; message_color_type=&#8221;uncode-solid&#8221; message_color_solid=&#8221;rgba(91,210,234,0.68)&#8221;]<\/p>\n<h1>Australian pine<\/h1>\n<p>[\/vc_message][uncode_list larger=&#8221;small&#8221; icon=&#8221;fa fa-tree&#8221; icon_color=&#8221;color-xsdn&#8221; css_animation=&#8221;top-t-bottom&#8221; animation_delay=&#8221;800&#8243; uncode_shortcode_id=&#8221;169614&#8243; icon_color_type=&#8221;uncode-palette&#8221;]<\/p>\n<ul>\n<li style=\"list-style-type: none;\">\n<ul>\n<li>Scientific name: <strong><em> Casuarina equisetifolia<\/em> L.<\/strong><\/li>\n<li>Common name: <strong>Casuarina, Australian pine, Horsetail Casuarina, Whistling Pine<\/strong><\/li>\n<li>Order: <strong>Fagales<\/strong><\/li>\n<li>Family: <strong>Casuarinaceae<\/strong><\/li>\n<li>Genus: <em><strong>Casuarina<\/strong><\/em><\/li>\n<li>Endemic: <strong>Non endemic species<\/strong><\/li>\n<li>Origin: <strong>Southeast Asia, Australia, Polynesia<\/strong><\/li>\n<li>Global distribution: <strong>The species exhibits a wide distribution in the American continents, Africa, coastal Mediterranean countries, as well as in South and East Asia<\/strong><\/li>\n<li>Distribution in Greece: <strong>Ionian Sea, coastal areas of Sterea Ellada<\/strong><\/li>\n<li>Altitudinal range: <strong>0-1,500m (in Colombia there are records of a maximum altitude of 2,650 m)<\/strong><\/li>\n<\/ul>\n<\/li>\n<\/ul>\n<p>[\/uncode_list][\/vc_column][\/vc_row][vc_row row_height_percent=&#8221;0&#8243; override_padding=&#8221;yes&#8221; h_padding=&#8221;0&#8243; top_padding=&#8221;0&#8243; bottom_padding=&#8221;0&#8243; back_color=&#8221;color-xsdn&#8221; overlay_alpha=&#8221;100&#8243; equal_height=&#8221;yes&#8221; gutter_size=&#8221;0&#8243; column_width_percent=&#8221;100&#8243; shift_y=&#8221;0&#8243; z_index=&#8221;0&#8243; style=&#8221;inherited&#8221; row_name=&#8221;infos&#8221; uncode_shortcode_id=&#8221;143868&#8243; back_color_type=&#8221;uncode-palette&#8221;][vc_column column_width_percent=&#8221;100&#8243; position_vertical=&#8221;middle&#8221; align_horizontal=&#8221;align_center&#8221; gutter_size=&#8221;3&#8243; override_padding=&#8221;yes&#8221; column_padding=&#8221;2&#8243; overlay_alpha=&#8221;100&#8243; shift_x=&#8221;0&#8243; shift_y=&#8221;0&#8243; shift_y_down=&#8221;0&#8243; z_index=&#8221;0&#8243; medium_width=&#8221;4&#8243; mobile_width=&#8221;0&#8243; css_animation=&#8221;zoom-in&#8221; zoom_width=&#8221;0&#8243; zoom_height=&#8221;0&#8243; width=&#8221;1\/1&#8243; uncode_shortcode_id=&#8221;965594&#8243;][vc_empty_space empty_h=&#8221;2&#8243;][vc_accordion no_toggle=&#8221;yes&#8221; typography=&#8221;advanced&#8221; sign=&#8221;plus&#8221; sign_size=&#8221;sm&#8221; label_border=&#8221;yes&#8221; content_border=&#8221;yes&#8221; gutter_simple=&#8221;0&#8243; titles_font=&#8221;font-804447&#8243; titles_size=&#8221;fontsize-922434&#8243; titles_weight=&#8221;700&#8243; titles_height=&#8221;fontheight-161249&#8243; active_tab=&#8221;0&#8243; uncode_shortcode_id=&#8221;924928&#8243;][vc_accordion_tab gutter_size=&#8221;0&#8243; column_padding=&#8221;0&#8243; title=&#8221;Ecosystem&#8221; tab_id=&#8221;1771584718601-2-81771656793597&#8243; slug=&#8221;ecosystem&#8221;][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;188831&#8243;]<\/p>\n<p style=\"text-align: justify;\">A species that thrives in tropical or subtropical sandy coastal areas and in semi-arid to slightly humid climates. It can grow in any type of soil, while sunlight is essential for its growth. It does not survive in low temperatures.<\/p>\n<p>[\/vc_column_text][vc_empty_space empty_h=&#8221;2&#8243;][\/vc_accordion_tab][vc_accordion_tab gutter_size=&#8221;0&#8243; column_padding=&#8221;0&#8243; title=&#8221;Botanical description&#8221; tab_id=&#8221;1771591735451-3-81771656793597&#8243; slug=&#8221;description&#8221;][vc_row_inner][vc_column_inner width=&#8221;1\/1&#8243;][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;907434&#8243;]<\/p>\n<p style=\"text-align: center;\">(height, leaf arrangement, leaf shape, root system)<\/p>\n<p style=\"text-align: justify;\">Fast-growing evergreen tree reaching a height of 10-30 m. Its crown is broad and pyramidal, with a final diameter of 8-10 m, while the trunk has a rough texture and a grey-brown or dark brown color. The leaves are dark green, simple, needle-like, and grow in whorls of 7 or 8. The root system is particularly extensive, a feature of the species that has led to its planting in areas where there is a need to limit soil erosion.<\/p>\n<p>[\/vc_column_text][\/vc_column_inner][\/vc_row_inner][vc_empty_space empty_h=&#8221;2&#8243;][\/vc_accordion_tab][vc_accordion_tab gutter_size=&#8221;0&#8243; column_padding=&#8221;0&#8243; title=&#8221;Reproduction&#8221; tab_id=&#8221;1771591801014-4-11771656793597&#8243; slug=&#8221;reproduction&#8221;][vc_row_inner][vc_column_inner width=&#8221;1\/1&#8243;][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;593378&#8243;]<\/p>\n<p style=\"text-align: center;\">(flowering period, flowers, seeds)<\/p>\n<p style=\"text-align: justify;\">Monoecious species with separate male and female reddish flowers. The male flowers are organized in spikes 1-4 cm long, while the female ones are organized in racemose inflorescences of 3-5 cm. The flowering period lasts from April through June. After fertilisation, woody, oval-shaped fruits develop. These fruits are 1-2.5 cm long and 1 cm in diameter, resembling cones. They contain seeds named samaras, with a pale brown hue, elliptical shape, and length of 5-8 mm, including their wings.<\/p>\n<p>[\/vc_column_text][\/vc_column_inner][\/vc_row_inner][vc_empty_space empty_h=&#8221;2&#8243;][\/vc_accordion_tab][vc_accordion_tab gutter_size=&#8221;0&#8243; column_padding=&#8221;0&#8243; title=&#8221;Noteworthy characteristics&#8221; tab_id=&#8221;1771591901860-5-01771656793597&#8243; slug=&#8221;characteristics&#8221;][vc_row_inner][vc_column_inner width=&#8221;1\/1&#8243;][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;114622&#8243;]<\/p>\n<p style=\"text-align: justify;\">Even though Casuarina equisetifolia resembles a pine due to the structure of its shoots and leaves, it does not belong to the conifers.<\/p>\n<p style=\"text-align: justify;\">Structures called nodules are observed on the roots of the trees, which are created by bacteria of the genus Frankia. Symbiotic relationships prevail between the two types of organisms, as the bacteria convert atmospheric nitrogen into forms that can be assimilated by the plant organism, allowing it to grow in nutrient-poor soils.<\/p>\n<p>[\/vc_column_text][\/vc_column_inner][\/vc_row_inner][vc_empty_space empty_h=&#8221;2&#8243;][\/vc_accordion_tab][vc_accordion_tab gutter_size=&#8221;0&#8243; column_padding=&#8221;0&#8243; title=&#8221;Historical information&#8221; tab_id=&#8221;1669205767-1-841771656793597&#8243; slug=&#8221;history&#8221;][vc_row_inner][vc_column_inner column_width_percent=&#8221;100&#8243; gutter_size=&#8221;3&#8243; overlay_alpha=&#8221;50&#8243; shift_x=&#8221;0&#8243; shift_y=&#8221;0&#8243; shift_y_down=&#8221;0&#8243; z_index=&#8221;0&#8243; medium_width=&#8221;0&#8243; mobile_width=&#8221;0&#8243; width=&#8221;1\/1&#8243; uncode_shortcode_id=&#8221;151656&#8243;][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;242506&#8243;]<\/p>\n<p style=\"text-align: justify;\">The first official description of the species was made in 1759 by Carl Linnaeus in his work Amoenitates Academicae. Its name is linked to the Cassowary (tropical bird) due to the resemblance of the foliage to its plumage. The specific epithet equisetifolia translates as &#8220;horsehair&#8221; and concerns the appearance of the leaves. In 1898, seeds of the species were introduced to Florida to be planted on the banks of canals and to stabilize the soil.<\/p>\n<p>[\/vc_column_text][\/vc_column_inner][\/vc_row_inner][vc_empty_space empty_h=&#8221;2&#8243;][\/vc_accordion_tab][vc_accordion_tab gutter_size=&#8221;0&#8243; column_padding=&#8221;0&#8243; title=&#8221;Uses&#8221; tab_id=&#8221;1669206843531-5-01771656793597&#8243; slug=&#8221;uses&#8221;][vc_row_inner][vc_column_inner width=&#8221;1\/1&#8243;][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;714931&#8243;]<\/p>\n<p style=\"text-align: justify;\">In the traditional culture of the Aborigines of Australia, the casuarina was used for the construction of weapons (boomerangs). The wood of the trees is hard and durable, traits that make it ideal for use in building constructions. It\u2019s also used as fuel and for paper pulp production. Furthermore, the bark\u2019s high tannin content makes it useful for dyeing leather and other fabrics. Various parts of the plant exhibit pharmaceutical actions, among which is antibacterial and anti-inflammatory activity. As it is resistant to salt and drought, the Casuarina grows even in sandy, coastal soils and in nitrogen-poor soils, while its roots stabilize the ground, making it useful in reforestation and erosion control projects.<\/p>\n<p><em><span style=\"color: #ff0000;\"><strong>(Note: Ethnobotanical data regarding the medicinal uses of plants must be handled with caution, and their application should be carried out exclusively under medical supervision)<\/strong><\/span><\/em>.[\/vc_column_text][\/vc_column_inner][\/vc_row_inner][vc_empty_space empty_h=&#8221;2&#8243;][\/vc_accordion_tab][vc_accordion_tab gutter_size=&#8221;0&#8243; column_padding=&#8221;0&#8243; title=&#8221;References&#8221; tab_id=&#8221;1771600746693-6-41771656793597&#8243; slug=&#8221;references&#8221;][vc_row_inner][vc_column_inner width=&#8221;1\/1&#8243;][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;324032&#8243; el_class=&#8221;micro-text&#8221;]<\/p>\n<ul>\n<li><a href=\"https:\/\/prosea.prota4u.org\/view.aspx?id=2993\">https:\/\/prosea.prota4u.org<\/a><\/li>\n<li><a href=\"https:\/\/powo.science.kew.org\/taxon\/urn:lsid:ipni.org:names:159856-1\/general-information\">https:\/\/powo.science.kew.org<\/a><\/li>\n<li><a href=\"https:\/\/www.worldfloraonline.org\/taxon\/wfo-0000590663\">https:\/\/www.worldfloraonline.org<\/a><\/li>\n<li><a href=\"https:\/\/www.researchgate.net\/figure\/Worldwide-distribution-of-Casurina-equisetifolia-Green-native-distribution-Purple_fig2_350449761\">https:\/\/www.researchgate.net<\/a><\/li>\n<li><a href=\"https:\/\/www.treesandshrubsonline.org\/articles\/casuarina\/casuarina-equisetifolia\/#:~:text=Casuarina%20equisetifolia%20is%20widely%20introduced,USDA%20Hardiness%20Zone%209\u201310\">https:\/\/www.treesandshrubsonline.org<\/a><\/li>\n<li>Maugham, W. S. (1928). The Casuarina Tree (Dust jacket illustration by W. E. Lefferts). Heinemann; George H. Doran Company.<\/li>\n<\/ul>\n<p>[\/vc_column_text][\/vc_column_inner][\/vc_row_inner][\/vc_accordion_tab][\/vc_accordion][vc_empty_space empty_h=&#8221;2&#8243;][\/vc_column][\/vc_row][vc_row unlock_row_content=&#8221;yes&#8221; row_height_percent=&#8221;0&#8243; override_padding=&#8221;yes&#8221; h_padding=&#8221;0&#8243; top_padding=&#8221;2&#8243; bottom_padding=&#8221;0&#8243; overlay_alpha=&#8221;100&#8243; equal_height=&#8221;yes&#8221; gutter_size=&#8221;0&#8243; column_width_percent=&#8221;100&#8243; shift_y=&#8221;0&#8243; z_index=&#8221;0&#8243; style=&#8221;inherited&#8221; uncode_shortcode_id=&#8221;877269&#8243; back_color_type=&#8221;uncode-solid&#8221; back_color_solid=&#8221;#ffeece&#8221; row_name=&#8221;epoxes&#8221;][vc_column column_width_percent=&#8221;100&#8243; position_vertical=&#8221;middle&#8221; align_horizontal=&#8221;align_center&#8221; gutter_size=&#8221;3&#8243; override_padding=&#8221;yes&#8221; column_padding=&#8221;0&#8243; overlay_alpha=&#8221;100&#8243; shift_x=&#8221;0&#8243; shift_y=&#8221;0&#8243; shift_y_down=&#8221;0&#8243; z_index=&#8221;0&#8243; medium_width=&#8221;4&#8243; mobile_width=&#8221;0&#8243; zoom_width=&#8221;0&#8243; zoom_height=&#8221;0&#8243; width=&#8221;1\/1&#8243; uncode_shortcode_id=&#8221;332089&#8243;][vc_custom_heading text_color=&#8221;color-jevc&#8221; text_font=&#8221;font-804447&#8243; text_size=&#8221;fontsize-922434&#8243; text_space=&#8221;fontspace-781688&#8243; uncode_shortcode_id=&#8221;120612&#8243; text_color_type=&#8221;uncode-palette&#8221;]During each season you will see:[\/vc_custom_heading][vc_gallery el_id=&#8221;gallery-2&#8243; type=&#8221;css_grid&#8221; medias=&#8221;86167,86169,86171,86173&#8243; explode_albums=&#8221;yes&#8221; screen_lg_items=&#8221;3&#8243; screen_lg_breakpoint=&#8221;1000&#8243; screen_md_items=&#8221;2&#8243; screen_md_breakpoint=&#8221;600&#8243; screen_sm_items=&#8221;2&#8243; screen_sm_breakpoint=&#8221;480&#8243; gutter_size=&#8221;0&#8243; css_grid_v_align=&#8221;bottom&#8221; css_grid_equal_height=&#8221;yes&#8221; media_items=&#8221;media|lightbox|original,title,caption&#8221; css_grid_images_size=&#8221;nine-sixteen&#8221; single_back_color=&#8221;color-rgdb&#8221; single_overlay_color=&#8221;color-jevc&#8221; single_overlay_coloration=&#8221;bottom_gradient&#8221; single_overlay_blend=&#8221;multiply&#8221; single_overlay_opacity=&#8221;70&#8243; single_overlay_visible=&#8221;yes&#8221; single_overlay_anim=&#8221;no&#8221; single_text_visible=&#8221;yes&#8221; single_text_anim=&#8221;no&#8221; single_image_anim=&#8221;no&#8221; single_h_align=&#8221;center&#8221; single_v_position=&#8221;bottom&#8221; single_padding=&#8221;2&#8243; single_title_dimension=&#8221;h4&#8243; single_border=&#8221;yes&#8221; single_css_animation=&#8221;alpha-anim&#8221; lbox_title=&#8221;yes&#8221; lbox_caption=&#8221;yes&#8221; no_double_tap=&#8221;yes&#8221; uncode_shortcode_id=&#8221;152816&#8243;][\/vc_column][\/vc_row][vc_row row_height_percent=&#8221;0&#8243; override_padding=&#8221;yes&#8221; h_padding=&#8221;0&#8243; top_padding=&#8221;2&#8243; bottom_padding=&#8221;2&#8243; back_image=&#8221;86162&#8243; overlay_color=&#8221;color-125592&#8243; overlay_alpha=&#8221;70&#8243; overlay_color_blend=&#8221;multiply&#8221; equal_height=&#8221;yes&#8221; gutter_size=&#8221;0&#8243; column_width_percent=&#8221;100&#8243; shift_y=&#8221;0&#8243; z_index=&#8221;0&#8243; style=&#8221;inherited&#8221; row_name=&#8221;texni&#8221; uncode_shortcode_id=&#8221;967188&#8243; overlay_color_type=&#8221;uncode-palette&#8221;][vc_column column_width_percent=&#8221;100&#8243; position_vertical=&#8221;middle&#8221; align_horizontal=&#8221;align_center&#8221; gutter_size=&#8221;3&#8243; override_padding=&#8221;yes&#8221; column_padding=&#8221;2&#8243; style=&#8221;dark&#8221; overlay_alpha=&#8221;100&#8243; shift_x=&#8221;0&#8243; shift_y=&#8221;0&#8243; shift_y_down=&#8221;0&#8243; z_index=&#8221;0&#8243; medium_width=&#8221;4&#8243; mobile_width=&#8221;0&#8243; zoom_width=&#8221;0&#8243; zoom_height=&#8221;0&#8243; width=&#8221;1\/1&#8243; uncode_shortcode_id=&#8221;129785&#8243;][vc_custom_heading text_color=&#8221;color-xsdn&#8221; text_font=&#8221;font-804447&#8243; text_size=&#8221;fontsize-922434&#8243; text_space=&#8221;fontspace-781688&#8243; sub_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;614274&#8243; text_color_type=&#8221;uncode-palette&#8221;]References in \u0391rt[\/vc_custom_heading][vc_column_text text_lead=&#8221;small&#8221; uncode_shortcode_id=&#8221;692532&#8243;]<\/p>\n<p style=\"text-align: justify;\">The species has been referenced in poetry, notably in the poem \u201cOur Casuarina Tree\u201d (1881) by Toru Dutt, where the tree serves as a symbol of her childhood and the nostalgia she feels. In W. Somerset Maugham\u2019s short story collection \u201cThe Casuarina Tree\u201d (1926), the tree symbolizes the English settlers in the Malay Peninsula and Borneo, who arrived after the pioneers and brought organization and stability to society, much like these trees stabilize the soil.<\/p>\n<p>[\/vc_column_text][vc_gallery el_id=&#8221;gallery-1&#8243; type=&#8221;carousel&#8221; medias=&#8221;86175,86177&#8243; carousel_lg=&#8221;3&#8243; carousel_md=&#8221;3&#8243; carousel_sm=&#8221;1&#8243; gutter_size=&#8221;0&#8243; media_items=&#8221;media|lightbox|original,caption&#8221; carousel_v_align=&#8221;middle&#8221; carousel_interval=&#8221;3000&#8243; carousel_navspeed=&#8221;400&#8243; carousel_nav=&#8221;yes&#8221; carousel_nav_mobile=&#8221;yes&#8221; carousel_nav_skin=&#8221;dark&#8221; stage_padding=&#8221;0&#8243; single_back_color=&#8221;color-rgdb&#8221; single_overlay_color=&#8221;color-jevc&#8221; single_overlay_coloration=&#8221;bottom_gradient&#8221; 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